#dentaneosuchus

2023 has been a year with its ups and downs, but one consistent thing has been its fossil crocs at least, giving us another interesting variety of species and genera not known before. Like last year, I'll go down each of them (including phytosaurs) and give you some key notes. And since I've done my best to make individual posts for them I'll link those when available. I'll also try to give translations where possible, but do note that sometimes I might switch around a word due to it just sounding awkward otherwise.

Scolotosuchus

Starting us off is Scolotosuchus basileus ("royal scythian crocodile"), a basal, rauisuchian-like animal from the early stages of the Triassic. An animal roughly 3 meters long, it is primarily known from vertebrae and some other material, which however does have interesting implications. Based on the anatomy of the vertebrae, it has been speculated that Scolotosuchus performed a lot of rapid movement of the neck, presumably while hunting. Furthermore, it might be that Scolotosuchus lacked osteoderms, instead having developed a bracing system for its body much more like that of dinosaurs and mammals. Artwork by @knuppitalism-with-ue (he'll pop up quite a lot)

Colossosuchus

Fast forward to the Late Triassic, the hayday of early Pseudosuchians, and we find our first phytosaur of the list. Colossosuchus techniensis ("colossal crocodile of the Indian Institute of Technology") was an enormous animal from India's Tiki Formation. This one is known from undisputably better material, perhaps some of the best on this list as we have an entire bone bed of these guys, possibly representing a mass death site. Size estimates of the large specimens generally range from 6 to 9 meters in length, also making it the largest animal of this list, with the authors favoring an estimated 8 meters for the largest individual. Of course this is all subject to change, as we don't know the precise proportions (the downside of a bone bed is that all the bones are kinda jumbled together and god knows what belongs to what). Artwork by Joschua Knüppe again, a female being courted by an overconfident and confused Volcanosuchus (told you so)

Kryphioparma

Ok I won't bother you with this one too much. Kryphioparma caerula ("blue mysterious shield") is an aetosaur from the Late Triassic Chinle Formation of Arizona. Now aetosaurs are cool of course, but the thing is that Kryphioparma is known from exclusively osteoderms, their large armour plates. While thats valid, its also not really exciting (as you could have guessed from the absolute lack of artwork). Best I can say is that it coexisted with a bunch of other aetosaurs, which surely would have been a sight to see back in the day.

Venkatasuchus

Now Venkatasuchus armatum ("Venkata's armoured crocodile") is in a similar boat. This Indian aetosaur too is only known from osteoderms, tho at least a lot more of them that were found together, meaning we got a much better idea of its shape. There are some interesting implications it has on paleogeography and how different animal groups spread across Pangea, but that's probably beyond the scope of this post. There is at least some art of it tho, including one piece by Joschua featuring Jaklapallisaurus, an early sauropodomorph.

Mystriosuchus alleroq

Now we got our first new species of a preexisting genus. While I did my best to make posts on new genera, which typically went hand in hand with their respective wikipedia pages, species are a different matter since they'd require me to overhaul and research the whole genus. Which for Mystriosuchus would go all the way back to 1896.

Anyhow, Mystriosuchus alleroq ("jawbone spoon crocodile") is the newest in a long line of Mystriosuchus species and has been recovered from western Greenland of all places. Like Colossosuchus it is known from multiple individuals, at least four in fact, And like Venkatasuchus, it appears to have wideraning implications for the spread of archosaurs during the Triassic. Oh look its Josch's art again, neat.

Jupijkam

The final phytosaur and the final Triassic taxon of the year, Jupijkam paleofluvialis ("horned serpent of ancient rivers") was recovered from Nova Scotia Canada and represents one of the northernmost phytosaurs (alongside obviously our friend Mystriosuchus alleroq above). It is far less complete than the other phytosaurs I covered so far, but still nothing to scoff at. It's also one of the youngest known phytosaurs, possibly having lived during the last stage of the Triassic. There's no art but here's an image of its snout from the original paper.

Turnersuchus

And with this we have reached the Jurassic and somewhat of a double feature. Ordering these via chronology makes this almost like storytelling. Turnersuchus hingleyae ("Hingley's and Turner's crocodile") is the oldest and basalmost named thalattosuchian, a group of crocodylomorphs that took to the seas. As such it shows a mosaic of features, already having begun to reduce its limbs, having a moderately long snout and still bearing osteoderms, which some of its descendents would come to lose. Turnersuchus dates to the Pliensbachian, a stage of the early Jurassic, tho the paper mentions how its not likely to hold its title as oldest member of its group for long as even older material has been discovered and is awaiting publication. Artwork by Júlia d’Oliveira and Joschua Knüppe

Torvoneustes jurensis

While Turnersuchus gives us a glimpse at what the earliest thalattosuchians were like, Torvoneustes jurensis ("savage swimmer of the Jura Mountains") represents them at one of their most derived. At 4 meters long, it must have been an impressive creature, entirely smooth and lacking osteoderms, highly reduced arms and a tail fluke to boot. There are some interesting features, as unlike other species of Torvoneustes, this one does not appear to have been as durophagous, instead appearing to be more of a generalist. It's the last Jurassic croc of the year and practically lived at the opposite end of this time period relative to our previous thalattosuchian, during the Kimmeridgian. Artwork by Sophie De Sousa Oliveira, not to be confused with Jùlia who drew Turnersuchus.

Comahuesuchus bonapartei

From marine to terrestrial, Comahuesuchus bonapartei is a member of the Notosuchia, land-dwelling cousins to the Neosuchia. This new species is known from the Cretaceous of Argentina and was described on the basis of a lower jaw, which differs from the previously described species of Comahuesuchus by the fact that the teeth are situated in individual sockets. Its jaws were short and wide, being described as U-shaped, giving it a somewhat pug-nosed appearance. In case you're wondering why I don't have the translation of the name thats because I can't access Bonaparte's first paper, so all I can say is that the new species is named after famed Argentinian paleontologist Jóse Fernando Bonaparte. Artwork by I mean you probably guessed it, its Josch again.

Aphaurosuchus kaiju

I have some mixed feelings on Aphaurosuchus kaiju ("powerless kaiju crocodile"), not just because I have to rewrite its genus wikipedia page that I am no longer happy with. No, Aphaurosuchus is a great genus in my opinion, given the complete nature of the holotype. But I do think that the species name of this second form is a bit of an exaggeration, seeing as it is to my knowledge not that exceptionally big. So why name it kaiju other than to sound cool? I also think it just kinda becoms funny when you consider the meaning of the genus name. Anyhow, it does look pretty mean, but thats to be expected from a baurusuchid. It lived during the Late Cretaceous in Brazil and the phylogenetic analysis that accompanied its description had some interesting implications that I'm curious to see tackled in the future. Other than that it's just another baurusuchid from a place crawling with baurusuchids, which tbh is pretty cool itself but doesn't really help make it stand out. Arwork by Paula Zeinner

Dentaneosuchus

It is the middle Eocene. France is entirely occupied by mammals. Well, not entirely... One indomitable reptile still holds out against the invaders. Yes, I am of course talking about Dentaneosuchus ("frightfull crocodile"), the titan of the Eocene, the terror of the jungle. Easily among the most exciting finds, this animal was an enormous sebecid previously thought to be a member of the genus Iberosuchus, but recent research not only shows its distinct but enormous. With a lower jaw around 90 cm long, it rivals even the famed Barinasuchus of Miocene South America. The total body lenght is of course a matter of debate given how little we know about sebecid proportions, with the paper lowballing it at 3 to 4 meters and I personally recovering something more akin to 5 meters. Regardless of the specifics, this would make it easily one of the top predators of its time, tho sadly this was not meant to last. It was among the last non-mammalian apex predators of Europe, before climate change and competition from mammals eventually drove them to extinction. Arwork by Joschua Knüppe and @mariolanzas

Baru iylwenpeny

Our last three entries were all members of the Notosuchia, the terrestrial crocodiles that existed from the Jurassic to the Miocene, so lets jump to the other major group of the time, the Neosuchians, specifically their most recent branch, the Crocodilians. Yes finally we are getting to crocs in the stricter sense.

Baru iylwenpeny ("divine crocodile thats good at hunting") is an animal we also knew for a while under the informal name Alcoota Baru based on where it was found. This year I spent a lot of time writing in the Mekosuchinae, Australasian crocodiles of the Cenozoic and just when I had finished the page for Baru, this guy got published. It's the largest, most robust and most recent member of the genus. Good at hunting is a good choice for the species name, as this animal was over 4 meters long with a head commonly likened to a cleaver and a hunting style speculated to consist of inflicting a lot of trauma on its prey with its massive teeth. I could gush about Baru for ages, but thats the important parts. It lived in central Australia during the Miocene only 8 million years ago. Artwork by the incredible @manusuchus

Antecrocodylus

Now for the last crocodilian described this year and one I remised to make a dedicated post for due to time constraints (I had a busy few months). Antecrocodylus chiangmuanensis ("before Crocodylus from Chiang Muan") is a close relative of the modern dwarf and true crocodiles from the Miocene of Thailand. It is only known from the back of the skull and an associated lower jaw, but it serves to highlight how little we know about the crocodiless of eastern Asia during the Miocene, which is a shame given that this region is crucial to deciphering where true crocodiles originated.

Alligator munensis

And our final taxon is Alligator munensis ("Mun river alligator"), which, obviously, is a member of the modern genus Alligator and thus most closely related to the still extant Chinese and American Alligator. Likely having been split from its Chinese relatives when the Tibetan plateau was lifted up, this species lived during the middle Pleistocene in Thailand and may have survived until the Holocene. It's head was short and robust and its teeth globular, which could indicate that it fed on hard-shelled prey like clams and snails. It was also small, likely not much bigger than 1.5 meters. Artwork by Joschua Knüppe

Issel Formation Diorama poster

An illustrated diorama depicting a scene out of the Issel Formation, dating back to Middle Eocene France. It features the sebecid Dentaneosuchus.

252mya.com/issel

Day 21: Dentaneosuchus

MAJOR NEWS FOR CROCODILE FANS In a newly published paper, Martin et al. describe a new genus of sebecid, Dentaneosuchus from the middle Eocene of France Whats cool about this animal is not just that its part of a group of terrestrial crocodylomorphs, i.e. croc relatives that lived and hunted on land But its also enormous The lower jaw was around 90cm long in the smaller specimen, which puts it in a similar size range as the enormous Barinasuchus from Miocene of South America pictured below (comparisson by Aledgn) this makes it perhaps the biggest land predator of Europe since the extinction of the dinosaurs.