Reconstructing Gigantopithecus blacki (WIP)
(Coming Soon- Finished Personal Reconstruction of a Gigantopithecus male, in replacement of Skeletal front page which is outdated (refer to Skull Anatomy))
Gigantopithecus blacki (Black's Giant Ape) [von Koenigswald, 1935] is the largest known ape and generally primate to exist, although many mysteries surround it for what it looks like and how it was in life. Some of these aspects of Gigantopithecus require some filling in of details with speculation from what is known and from extant and extinct relatives. For a lot of the things that will be discussed here, there is a lot of room for speculation and other forms of decision.
In this post, we will cover multiple topics in order: Phylogenetics, skeletal anatomy, size, skull anatomy, soft tissue, musculature, posture, mobility, integument, coloration, diet, social behavior, and sexual dimorphism. Most of these sections will be accompanied by charts that represent the topics, reference images, and examples.
As with my Nanuqsaurus post, Gigantopithecus is insanely fragmentary and much of it is open to interpretation. Some of the information here isnât set in stone since Paleontology is an evolving field with new discoveries. Information may be eventually outdated and updated, though if anything is outdated, wrong, or not clear, feel free to point out these mistakes, this includes citations. This page will be actively updated.
People also have different interpretations of Gigantopithecus reconstructions and this page aims to help describe the core features that Gigantopithecus likely had while toying around with speculative structures, alternate hypotheses, etc in some sections.
Also as of initially publishing this, some things may be somewhat incomplete. Sections regarding the Ecology and Range of Gigantopithecus are for now missing mainly because it isnât a high priority and some charts might be missing. This will be added later.
Ponginae (10-15 mya split)
>Gigantopithecus (10.14 mya split, lived 2-0.3 mya)
>>Lufengpithecus (lived 6.2 mya)
>>>Ankarapithecus (lived 5-7 mya)
>>>>Sivapithecus (lived 12.5 to 8.5 mya)
The relationship between Gigantopithecus and other primates regarding how close they are related is one of the important things to think of when reconstructing the ape, as it would be impactful to a degree to take into consideration closely related species. Throughout history, there has been debate where it should be placed. It originally was viewed as a hominid, closer to us than other primates, although that would be disproved (Broom, 1939). What would be found is that Gigantopithecus would fall under the Pongidae family, the lineage that includes the current three species of Orangutan. Ponginae overall has multiple extinct members and clades, Pongini, Sivapithecini, and Lufengpithini. The current findings indicate Gigantopithecus being more basal in the Pongine lineage, closer to Sivapithecus and Lufengpithicus. It is more specifically classified as a basal offshoot in the overall lineage (Zhang & Harrison, 2017).
Because Gigantopithecus is fairly basal in the Ponginae lineage, a lot of traits would be impacted from where it is placed. If it were directly sister to Orangutan with a more recent ancestral split, the âovergrown Orangutanâ âpaleomemeâ would be more fair, although Gigantopithecus is fairly distant from Orangutans. Gorillas split from us the same amount of time ago as Gigantopithecus split from Orangutans (Welker et al., 2019).
âThis results in a divergence time of GigantopithecusâPongo of 10.14 Mya (4.76-15.79 Mya, 95% HPD interval). The divergence of Gorilla from the Homo/Pan branch is estimated at 8.59 Mya (4.62-13.56 Mya, 95% HPD interval), and the divergence of Homo and Pan at 5.78 Mya (2.64-9.53 Mya, 95% HPD interval). These are largely consistent with, but somewhat younger than, previous estimates52,54, possibly due to a mutation slowdown on these lineages compared to the Pongo lineage, which is not taken into account here. However, they seem in agreement with the fossil record indicating the origin of hominins around 6-8 Mya and the dating of a possible early Gorillini (Chororapithecus) around 7-9 Mya54-58.â (Welker et al., 2019).
Gigantopithecus itself wouldâve likely been more derived in its own lineage for how far it lived into the Pleistocene to when it split from the Pongine lineage. What will be covered soon in [Skeletal Anatomy], a lot of the anatomical traits Gigantopithecus had would be different and similar to many extant apes and the phylogenetic classification should be taken into consideration when reconstructing this great ape.
Note - (Welker et al., 2019) is a great source of information regarding the placement of Gigantopithecus and dives into very technical explanations which canât be easily covered here. Feel free to give the paper a read to understand the actual reasoning behind the placement.
(Coming soon - Chart of different body interpretations)
One heavily debated aspect of Gigantopithecus is the sort of body structure it possesses. Many reconstructions imply a more Pongine look with the long forelimbs and shortened hindlimbs, while some imply more even and equally proportioned limbs more of a Gorilla. When looking at Gigantopithecus, it is easy to take from the absolute size of the animal, even if you go with lower estimates (covered in the size section), it still outweighs the largest Gorilla species Gorilla beringei graueri (Eastern Lowland Gorilla) in mass at an male adult size of 170 kg. Combined with the fact that the diet of Gigantopithecus includes mainly ground plants (covered in the diet section), there is no doubt that the animal is terrestrial when fully grown (Zhang & Harrison, 2017). Though it being terrestrial would affect the proportions of the limbs and robustness. Arboreal animals tend to have adaptations that allow them to be more suited for their lifestyle. Orangutans spend much of their lives in the trees and have heavily disproportionate limbs, long arms and short legs. While Orangutans may be the closest living relative to Gigantopithecus, it would be overall better to take into consideration Gigantopithecusâ differences from it. Being intensely terrestrial would have arm to leg proportions more similar to a Gorilla. Considerably built and near-equal in length. For actual proportional length to torso for if the limbs were longer or shorter, there as always isnât a true answer, though it would be most likely for the limbs to be at least the proportions of a Chimpanzee, as Gigantopithecus evolved from insanely arboreal ancestors in the Pongine lineage. The limbs being short proportionately generally wouldnât do any good for what Gigantopithecus generally ate since it was eating foods that were close to the ground and at head level.
For overall frame such as shoulder and torso width, great apes tend to have large bodies, though they vary in width and proportions. Gorillas have larger and wide shoulders and large hips (Zihlman et al., 2011), while others tend to have narrower shoulders or hips. This correlates to the overall mobility of the animal, as Gorillas are more robust to adapt to their more terrestrial lifestyle and mobility, while Orangutans and Chimpanzees are built the way they are mainly for arboreal adaptations. Gigantopithecus probably had something more in line between a Gorilla and Orangutan build, although likely leaning to Gorillas. Larger shoulders and hips are generally better for a terrestrial lifestyle since there isnât an evolutionary push for an orangutan build. Though this comes to say that with where Gigantopithecus is placed in classification as covered in [Phylogenetics], it is safe to conclude that while there would be some convergent traits and shared traits between the great apes Gigantopithecus would stand out differently drastically. Most of its anatomy would be adapted for what it ate, what it had to deal with in life, sexual dimorphism, and its terrestrial adaptations. When reconstructed in flesh, Gigantopithecus would be noticeably distinct and this should be done when reconstructing it. While stylistically in a fictional sense creative liberties can be taken if wanted, scientifically it shouldnât just be an overgrown Orangutan, but inspiration can be taken from Orangutans.
(Coming soon - Size Charts of minimum, conservative, maximum sizes)
One of the more controversial topics regarding Gigantopithecus, many options are available for size. Many people tend to downsize Gigantopithecus to an extreme and many people upsize it to an extreme. It is difficult to pinpoint which size Gigantopithecus wouldâve had because of the lack of material outside of teeth and mandibles. It depends on the scaling of the mandible to the skull along with the scaling depending on what proportions are being used. Multiple estimates use certain models for either Gorillas or Orangutans in which both drastically differ in frame and will yield different results, though as discussed before, the frame Gigantopithecus wouldâve had would be different from both.
The different sizes gathered range from 204 kg (Zhang & Harrison, 2017), 280kg (Zhang & Harrison, 2017), to 300 kg (Fleagle, 2013), Though directly quoted by the 2017 Zhang & Harrison paper, âWithout postcranial remains it is simply not possible to obtain a reliable estimate of the body mass for G. blacki, but 200â300 kg does seem like a reasonable guideâ (Zhang & Harrison, 2017). The paper goes on to say that the size would restrict any arboreal behaviors as discussed in [Skeletal Anatomy], though reliable size estimations can really only be gathered by postcrania since measurements based off mandibles and teeth can lead to improper results purely because of proportions. It is important to consider proportions of the primate of head to body, limbs to body, etc. A smaller head to body ratio would lead to a much larger primate, while a larger head would lead to a smaller primate, with this being amplified through measurement estimations based off Orangutans and Gorilla skeletons. Alternatively, limb proportions would also change. Longer limbs would mainly change the height of the estimations and only contribute very little to overall mass, though dimensional size is important to consider in some cases, especially for size charts.
âWithout postcranial remains it is simply not possible to obtain a reliable estimate of the body mass for G. blacki, but 200â300 kg does seem like a reasonable guide. Such a size (which overlaps with the upper end of the body mass range for extant male gorillas) may have precluded or greatly restricted arboreal behaviors in G. blacki, but once again postcranial remains are needed in order to determine its inferred locomotor repertoireâ (Zhang & Harrison, 2017).
Disclaimer- This skeletal is slightly out of date, some aspects will soon be changed of it (lineart smoothing, orbit size, side information)
Another debated aspect of Gigantopithecus is the facial and skull structure, mainly the robustness of the skull and the overall skeletal and soft-tissue structure of the entire skull. What is important to consider is the type of diet Gigantopithecus had (which is covered in the diets section), the size and body proportions, and the phylogenetics in relation to other close species. Facial anatomy tends to vary in ape species. Things like cheekbone width and size, sagittal crest size, how prominent the brow ridge is, size of the face in relation to the braincase (essentially, how much of the front of the face is forehead), orbit size, etc.
Gigantopithecus is well known for being primarily herbivorous, chewing tough fibrous plants such as grasses, low lying plantlife, fruits, and other low plantlife (as covered in diet). The thickness of the maxilla can be found from getting the proportional thickness of any of the four mandible pieces we have. The actual robustness of the skull can be compared to both Orangutans and Gorillas. Gorillas somewhat have a similar niche, being megafaunal terrestrial herbivores and eating similar things, mainly roots and leaves, Gigantopithecus has the same overlap with their diet. Tougher plants in diets tend to select for more robust facial features to allow for more musculature for the jaw. Primates have this very prominently, the aforementioned Gorillas and multiple species of non-ape primates also have robust jaws along with Orangutans. Larger cheekbones, overall a robust facial structure, and potentially large sagittal crests would help to accomplish the stronger bite force (Balolia et al., 2017). Sagittal crests would mainly be more prominent in males than females, mainly jaw clamping and bite force would be the functional use. The existence of sagittal crests have been linked with cranial growth and individual male success, with the crests being sexually dimorphic (Balolia et al., 2017). With the reduction of canines in males indicating lower use of canine display in male-male combat (Zhang & Harrison, 2017), bite force can likely be used as a substitution against other individuals to inflict damage. This works overall as a dietary advantage for eating tough foods, like fruits and tough plantlife, and as a general defensive function against predators if needed. Though with the existence of a large sagittal crest, this would amplify the temporal muscle size (Balolia et al., 2017). The actual size of the crests would be fairly large proportionately, since larger cranial sizes correlate with sagittal crest size. Gigantopithecus with their large skulls and for males would especially have this, especially for large males (Balolia et al., 2017). The topic of sagittal crests is further covered in the [Dimorphism] for speculative comparisons between male and females.
With all of this in mind, Gigantopithecus' face wouldâve been very robust in adaptation to what it was eating and size. Other things to consider is orbit size and while it may not fully impact external anatomy, it does still have an impact.
[Non-Musculature Soft Tissue]
Non-musculature soft tissue includes things like cheek flanges and throat pouches. While self explanatory, both structures are purely hypothetical when it comes to Gigantopithecus. The closest confirmed relatives that have this sort of structure are obviously the 3 Orangutans. Cheek flanges are dimorphic between both sexes with females lacking them and older males having them. Throat pouches exist between both genders and usually are for sound amplification and production (Utami, 2002). Since there was extreme dimorphism and competition between Gigantopithecus individuals, it would be safe to assume that there mightâve been some sort of structure, although this again is speculation with some reasonable basis. If the cheek flanges hypothesis is going to be done on Gigantopithecus, they likely wouldâve had them more reduced or modified compared to Orangutans, likely more thicker, rounder, etc.
[Musculature, Posture, and Mobility]
Gigantopithecus musculature would be like that of most other great apes. The important thing to take into account for musculature is how terrestrial Gigantopithecus was, and since Gigantopithecus has been concluded as being extremely terrestrial as opposed to being arboreal like Orangutans, musculature will likely differ from what would be normally expected from arboreal apes. While muscle groups would be most similar in structure to Orangutans, distribution would mostly likely be like Gorillas. Gorillas have higher hindlimb muscle composition compared to their forelimbs, while Orangutans are the opposite for different reasons. Gorillas bear much of their weight constantly on the ground being terrestrial creatures (while also being occasionally arboreal). Orangutans spend most of their times living an arboreal life, lifting their body weight up mainly with their arms (Zihlman et al., 2011).
The rest of this section will be written and covered later.
Existence of hair on Gigantopithecus is not up for debate, it does exist on it. Though the main question is the length and distribution of hair on Gigantopithecus. Many other primates, as in almost all, tend to lack hair around their faces, hands, feet, and occasionally their back thighs and chests. Multiple primates and especially Orangutans possess beard and mustache hair structures. As discussed in [Social Behavior and Dimorphism], beards and mustaches is often used as a sign of maturity, especially in males. It is impossible to rule out the existence of such facial hair structures as it is still highly unknown what Gigantopithecus wouldâve had, although it is likely that it did appear.
Other forms of hair âdisplayâ or functionality structures are common, such as a hypothetical sort of âmaneâ around the head, which some primates do feature. What wouldâve been likely to appear in life is the existence of long hair on the arms and legs, and potentially sides of the torso. Orangutans and Gorillas often have this shaggy hair structure on their lower half of their bodies, which is too also entirely plausible for Gigantopithecus.
Though what would be highly unlikely is a low amount of hair coverage, like elephants or humans on Gigantopithecus. The notion that Gigantopithecus should be mostly hairless or hairless which has been spread around recently is mainly from the reconstruction mentality of Archosaurs. Many of these arguments applied onto Gigantopithecus are the same arguments that work for animal groups like Dinosauria as a whole. Mammals should be treated differently compared to Archosaurs when it comes to reconstructions. The other issue that comes into play even for mammals is size. As discussed in the [Size] chapter, even the largest possible size estimation would still do very little to impacting fur distribution as many mammals of that size still possess a fair bit of hair. To conclude, Gigantopithecus despite its size wouldâve maintained a similar or the same amount of hair density and thickness as whatever is being modeled, from Orangutans, to Gorillas, to Chimpanzees.
The coloration debate is as always with any animal we donât have any preservation or records for. We can really only infer based on habitat, ecology, and anatomy. Animals that tend to live in tropical environments, like Gigantopithecus, have more varied coloration. Plenty of non-avian reptilians, aves, mammalians, amphibians all are often varied for different reasons. Primates too are often varied. Animals like Orangutans have orange coloration mainly for camouflage and a result of how their skin absorbs color. Orangutans tend to live in canopy trees and reside in more brown surroundings, like peat-colored landscapes, dead leaves, branches, etc. It is important to consider that both non-human mostly terrestrial apes, Gorillas and Chimpanzees, have black fur. This too mostly comes in use for camouflage blending in the forest floor.
Because Gigantopithecus is a terrestrial species with limited arboreality, it would be safe to infer that it had coloration more suited for a forest floor ecosystem. Darker coloration on the body fur, probably some countershade, etc.
Though coloration will always remain a debate and up for speculation, especially considering potential patches of display, like lightened shoulder fur being white. It is highly important to consider the habitat of Gigantopithecus so anything that would help for forest camouflage would be beneficial to survival.
This section is a Work in Progress, it will feature its ecological role and the range of where it lived.
The diet of Gigantopithecus is probably one of the most well understood things about the animal. With the huge collection of teeth, mandibles, and analysis and research of both. Multiple recent studies have shown that mainly leaves and occasionally fruits, overall C3 plants comprise their diet. For fruits, some indication exists for a partially fruit diet due to a Gigantopithecus population and some dissolving of enamel on found teeth. Their large and robust jaws wouldâve helped to come into use for chewing through harder or more fibrous foods, a lot of which would include ground-plants like roots, stems, leaves, and grasses, especially for Bamboo. While Gigantopithecus was not strictly an eater of bamboo as originally implied, it did comprise of their diet to a significant degree. Overall, they wouldâve been generalists with their food options more similar to Chimpanzees (Zhang & Harrison, 2017; Bocherens et al 2017 ).
Over the course of the Pleistocene, multiple events for forests expanding and shrinking occurred. This would directly impact Gigantopithecus feeding habits to adapt to the moment, with some evidence focusing on higher fruit consumption at certain periods of their existence.
As for meat consumption, nothing for full proof exists for the consumption of meat sources. Although what can be inferred from other primates is that insects like termites would have provided some supplementary substances when needed for extra protein. Gorillas and Orangutans often engage in termite-eating behavior despite being mainly herbivorous, Gorillas more-so herbivorous. Though carnivory of larger vertebrates or vertebrates wouldâve been very low. Gorillas rarely engage in the consumption of small vertebrates like rodents compared to their termite behavior, while active hunting is practically non-existent and same goes for scavenging. Gigantopithecus wouldâve likely fell into the same category and with intensive research on their diets through teeth examination and isotopic analysis, nothing comes back for intensive meat consumption. Though it is fair to speculate that very rare scavenging wouldâve occurred in times in need.
To add to overall feeding behavior, tool usage wouldâve likely appeared multiple times. Tool usage is a common behavior to many primates, especially great apes. As discussed with termites, âfishingâ wouldâve likely been used where a stick is used to gather termites and ants from bark, mounds, etc. Other tools probably came into play as well such as using rocks to smash open nuts as with Gorillas (Breuer et al., 2005). Orangutans too have been observed many times with tool use behavior as well, and tool usage tends to be very useful for diets.
[Social Behavior and Dimorphism]
(Coming soon - Size Chart of Male and Female Comparison)
The social behavior of Gigantopithecus has been debated for and against for how social the animal was in life, if it were like Orangutans with being solitary and territorial, Gorillas travelling in a male-dominated group, or Chimpanzees in a hierarchical tribe. What is true is that sexual dimorphism in Gigantopithecus exists, but enlarged canines were likely not part of the dimorphism, suggesting low usage of them in intraspecific fights and threat display (Zhang & Harrison, 2017). Likely, there wouldâve been other locations for possible display dimorphic features. While what exactly is unknown, it could likely boil down to possible fur coloration, fur length, prevalence of soft tissue, a behavioral display, sagittal crests, or muscular tissue. For many animals when it comes to this, it is hard to pinpoint what exactly would be the case. Though as for primates most times it does go for the things listed. Gorillas often use physical strength against eachother while Orangutans as well but with the addition of cheek flanges to signal maturity and Chimpanzees mainly differ in size and hierarchical status. It can be speculated (within reason at least) that Gignatopithecus can use any means of methods for dimorphism, though a direct copy-paste from Orangutans would be somewhat unlikely due to how highly variable primates can be as discussed in [Non-Musculature Soft Tissue].
There is an intense size difference between both males and females based off mandible sizes and what was described as both. Male mandibles and female mandibles (III and I) differ in size with the male mandible being 40% larger (Zhang & Harrison, 2017).
Males probably were more solitary than females or at least preferred to avoid other males by having territories to themselves, as with Gorillas, Orangutans, and Chimpanzees. Such dimorphic differences would indicate the male-male competition previously mentioned with the size difference (Zhang & Harrison, 2017). For actual societal structure many things still remain a mystery. We canât conclude if there were tribes with a leading female, female-only tribes, or all were solitary. With how different each primate is in behavior, there can be a lot of variation and differences in Gigantopithecus from other primates.
(Coming soon - Sagittal crest variation in males)
To quickly cover the concept of sagittal crests, â(sagittal) cresting in primates has been traditionally linked with the need for largerâbodied individuals to have sufficient attachment area for the temporalis muscle (Ashton & Zuckerman, 1956; Robinson, 1958; Holloway, 1962; Hofer, 1974). The reasoning is that because increases in the size of the braincase do not keep pace with increases in body size, there would not be enough space for the temporalis muscle in largerâbodied individuals without the additional surface area provided by sagittal crests (Robinson, 1958; Hofer, 1974) ... (the) presence of a sagittal crest increases the height of both the frontal and lateral profile of the head, and recent behavioural research suggests that the sagittal crest in G. g. gorilla males is associated with male reproductive success. Caillaud et al. (2008) found a positive relationship in breeding silverbacks between male sagittal crest size (measured as size of the adipose hump, using photographs of head profiles) and the number of females associated with that maleâ (Balolia et al., 2017). The role of sagittal crests wouldâve played a large role in sexual dimorphism, especially in competition. Variation between males would be as described, indicating the male-male competition and reproductive success.
Gigantopithecus is overall a mysterious fossil animal. Many of it remains a secret purely because of the lack of reliable information, that being skeletal remains. Though despite its fragmentary nature, much of it can be inferred from other relatives and primates, though this has to be taken seriously to consider certain traits of the primate. Itâs unreliable to directly copy off of a primate 1:1, and it is also a problem just adding random traits with little justification, some things have to have a reasoning rather than just slapping on a feature (or removing) with little justification. This will occasionally result in diving into these topics and researching them to understand.
Though I do hope many of you have learned from this reconstruction guide and found it informational. Many things of this gigantic ape is up to how you view and see it. Have fun with it.
Thanks for reading, and as always, feel free to contact or respond to this to correct, add, modify, or remove anything and what specifically. Have a great day.