Dictyonema glabratum

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Dictyonema glabratum

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Sticta with Dictyonema (Cora) by Richard Droker Via Flickr: cloud forest near Mindo, Ecuador need to go over these, e.g. compare 5th photo here, Lobariella www.facebook.com/photo.php?fbid=10153453250398371&set... regarding Dictyonema - apparently Cora back in Dictyonema, although wonder why Cora seems to still be used in recent papers www.facebook.com/photo.php?fbid=2882538018482531&set=... my lichen photos by genus - www.flickr.com/photos/29750062@N06/collections/7215762439... my photos arranged by subject, e.g. mountains - www.flickr.com/photos/29750062@N06/collections
Dictyonema glabratum - A lichen species that is actually hundreds of species
Also known as Cora pavonia, Dictyonema glabratum (Hygrophoraceae) is a tropical, cyanobacterial basidiolichen; it, means that is formed by the symbiosis between cyanobacteria and basidiomycete fungi (unlike most lichen which are formed by the symbiosis of a green algae and an ascomycete fungi).
Thallus of this lichen is large, up to 50 cm or more in diameter, with lobes of 2–10 cm wide, and the upper surface distinctly zonate, pale bluish to greenish gray. Dictyonema glabratum is distributed in the United States (Florida, Hawaii), southern to north-central Mexico, Central America, the Caribbean, South America, and Africa (Mauritius Island, E of Madagascar).
Dictyonema glabratum is an important component of threatened paramo ecosystems, where it acts as a biological fertilizer due to its ability to fix atmospheric nitrogen. But most surprising of this lichen is that a research recently published (21014) has demonstrated that D. glabratum, which was long believed to represent a single species, is actually at least 126 phylogenetically and morphologically distinct species, with statistical analysis predicting more than 400.
References: [1] - [2] - [3]
Photo credits: [Top: ©Daniel Winkler | Locality: Bolivian Amazon, 2013] - [Bottom: ©newmy51 (CC BY-NC-SA 2.0) | Locality: Coroico, Bolivia, 2008]
Liquen / Lichen | ©Jorge Restrepo
Unidentified species of lichen. Cocorna, Antioquia, Colombia.
Does anyone know the species?
Thanks to paisleytaxonomy, who has suggested an identification of this lichen as Dictyonema glabratum. Reviewing the literature it certainly seems that species (although the genus contains about 22 species).
The interesting thing is that Dictyonema is a symbiosis between a basidiomycete fungi and a scytonematoid cyanobacteria, making it both a basidiolichen and a cyanolichen, which is a very rare combination [1].
This very cool lichen, is one of the few true basidiolichens (closely related to true mushrooms as opposed to the vast majority of lichens which are related to things like cup fungi, morels, etc.). Apparently the relative quantity of algae to fungus can make a large difference in thallus structure. This particular specimen is dominated by the fungus. The hymenium, or spore-bearing surface, is smooth (hidden below). The alga is a cyanobacterium in the genus Scytonema [2].