Small white butterflies – the most common butterfly in this year’s British count, and one of the species that experienced a rise in numbers thanks to good weather.
Photograph: Will Langdon/Butterfly Conservat/PA

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Small white butterflies – the most common butterfly in this year’s British count, and one of the species that experienced a rise in numbers thanks to good weather.
Photograph: Will Langdon/Butterfly Conservat/PA

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A white wagtail on blossom in central China’s Henan province.
Photograph: Mei Yongcun/Xinhua/Barcroft Images
A black bear scratches his back in the Rocky Mountain wilderness, Canada
Photograph: Jill Cooper/Alamy
The vision by Jan van der Greef, The Netherlands – winner, Black and White
Perfectly balanced, its wings vibrating, its tail opening and closing, with its tiny feet touching the spike for just an instant, an eastern mountaineer hummingbird siphons nectar from the florets of a red-hot-poker plant. For a number of days Jan had observed the birds in the garden of his hotel in southern Peru. He noticed that an eastern mountaineer – a species found only in Peru– would rotate around the red-hot-poker as it fed. As it moved behind a spike and its tail closed, a beautiful cross appeared.
Photograph: Jan van der Greef/2018 Wildlife Photographer of the Year
Kuhirwa mourns her baby by Ricardo Núñez Montero, Spain – winner, Behaviour: mammals
Kuhirwa, a young female of the Nkuringo mountain gorilla family in Uganda’s Bwindi Impenetrable Forest, would not give up on her dead baby. What Ricardo first thought to be a bundle of roots turned out to be the tiny corpse. Guides told him that she had given birth during bad weather and the baby probably died of cold. At first Kuhirwa had cuddled and groomed the body, carrying it piggyback. Weeks later, she started to eat what was left of the corpse, behaviour the guide had only seen once before.
Photograph: Ricardo Núñez Montero/2018 Wildlife Photographer of the Year

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Bees on a sunflower in Heppenheim, Germany
Photograph: Armando Babani/EPA
A herd of American bison of genetically pure breed that have been reproduced as part of a bison and grassland conservation program at El Uno ranch in Chihuahua state, Mexico
Photograph: Herika Martinez/AFP/Getty Images
Mud-rolling mud-dauber by Georgina Steytler, Australia – winner, Behaviour: invertebrates
It was a hot summer day, and the waterhole at Walyormouring Nature Reserve, Western Australia, was buzzing. Georgina had got there early to photograph birds, but her attention was stolen by the industrious mud-dauber wasps. They were females, digging in the soft mud at the water’s edge, then rolling the mud into balls to create egg chambers for their nearby nests. A female builds her external nest completely out of mud, cylindrical chamber by chamber, which cement together as the mud hardens.
Photograph: Georgina Steytler/2018 Wildlife Photographer of the Year
A bird rests on the head of a white-tailed deer roaming free in San Jose Villanueva, El Salvador.
Photograph: Marvin Recinos/AFP/Getty Images
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A black bear scratches his back in the Rocky Mountain wilderness, Canada
Photograph: Jill Cooper/Alamy
Humpback whale near the Broughton Archipelago, Great Bear Rainforest, British Columbia, Canada.
Photograph: Alamy Stock Photo
The golden couple by Marsel van Oosten, The Netherlands – grand title winner, Animal portraits
A male Qinling golden snub-nosed monkey rests on a stone, joined by a female from his group. Both are watching an altercation down the valley between the lead males of two other groups in the 50-strong troop. It’s spring in the temperate forest of China’s Qinling mountains, the only place where these endangered monkeys live. To show both a male’s beautiful pelage and striking blue face, Marsel had to shoot at an angle from the back. It took many days observing the group to achieve his goal.
Photograph: Marsel van Oosten/2018 Wildlife Photographer of the Year
Barbary macaque monkeys play around in the Upper Rock area of the Gibraltar nature reserve. They are the only wild monkey population in the European continent
Photograph: Sean Gallup/Getty Images
A black or Yunnan snub-nosed monkey in Deqen prefecture, Tibet.
Photograph: Xinhua / Barcroft Images

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Monkey ball by Thomas Kokta (Germany)
Cold temperatures on Shodoshima Island, Japan, can lead to monkey balls, where a group of five or more snow monkeys huddle together to keep warm. Thomas observed a large group exhibiting this behaviour close to a tree, giving him the vantage point he needed. A few days and a tree-climb later, he got the image he wanted.
Photograph: Thomas Kokta/2016 Wildlife Photographer of the Year
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