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It is broadly the same point about learning from prediction errors that modern Pavlovians make and that Wolfram Schultz found manifest in the human dopamine system.
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Error Theory note
The brain updates internal representation of the environment by using the mismatch between the predicted state/outcome and the actual one, termed prediction-error. In parallel, time perception in the sub-second range is crucial for many behaviors such as movement, learning, memory, attention and speech. Both time-perception and prediction-errors are essential for everyday life function of an organism...  https://doi.org/10.1101/766204
