What if Plagg and Tikki never had dedicated holders before?
Like, what if there was a world where all the other Motherbox miraculous have been handed out and used before, but not the ladybug or black cat?
(TLDR: Eldritch god Tikki and Plagg love their strange squishy humans, and will crush anyone and anything that makes them cry.)
All of the other Kwami have experienced gaining, having, and losing human partners, but not Tikki and Plagg, as they have always been deemed too powerful and too dangerous to give to others, and they were rarely, if ever, used by the Guardians themselves, and if they were, it was never long enough to build any kind of connection. To the point that, if they ever give a kwagatama to anyone, they would be unable to contact anyone with it as they have never truly had past holders.
In a world like this, I like to imagine that either Master Fu was pushed to the edge of desperation by Hawkmoth and didn't see any other combination of Miraculous that could stop him while protecting Paris, OR, Plagg and Tikki came to that conclusion, and went out and chose wielders without permission.
Either way though, I think the results would end up the same.
Plagg and Tikki have never had a Human of their own before. They've only ever been able to here their sibling Kwami gush or whine about their holders. They had no idea what to expect, especially not from untrained children. But in the end, they learn the others were right: there is NOTHING quite like having your own human.
Plagg starts out trying to be aloof and disinterested. They are gods, after all, deigning to work with humans so as to not rip apart the fragile material world with their might. It would be silly to be anything less but flippant and relaxed like a deity before a worshiper.
So quickly does that fall apart once he's actually with Adrien. His human, no, his kitten, is the kindest creature he has ever met. And though he is very sheltered, he also far from naive, with a whip sharp mind and a protective streak a mile wide. But when it comes to himself, Plaggs chosen cannot lift a finger in self defense. He tucks his head and lets blows both metaphorical and literal rain down on him. This too smiley, too innocent, goody-two-shoes is the opposite of the unstoppable force of nature that Plagg represents.
So why is it that this small human child is the first being besides Tikki he has ever felt loyalty too? Why does his heart ache when the boy cries, and his anger surge when he sees how little Adrien's father cares for him? What has this human done to him?
Plagg doesn't know. Possibly can't know, even. But he knows he does. He knows this little blonde kit is his, and rules of the guardians be damned, he will do whatever it takes and whatever he wants to protect this little scrap of human and keep him happy.
Tikki, for her part, had gone into this new experience excited to meet her human. To finally interact closely with one of the most complicated lifeforms of this world she and Plagg had created together when they were still one being.
She had all these ideas for how she would shape the young champion. That she would create a paragon of virtue, a perfect shining hero. An example for all others for the proper way to act, to be. Not just as a miraculous holder, but as a human and sentient being. She went in fully expecting to follow every rule the Guardians had laid out, and to act as a wise and kind, but also impersonal and detached, mentor.
That plan survived barely two weeks with Marinette Dupain Cheng.
The poor thing was a kind, brave, and creative soul, who Tikki nearly crushed under the weight of her lofty expectations. The girl ran herself ragged trying to be what Tikki, and the order, wanted her to be. And it nearly broke her.
Tikki learned for the first time exactly how fragile human children can be, and how much pressure her human already put on herself. As the god of creation carefully helped the young designer recover and rebuild, she witnessed how few support systems she had, how little confidence she possessed, and how lonely and isolated she felt.
Tikki abandoned her lofty dreams of being a widened guide and teacher, her future plans of a perfect hero, for the simple task of being the lonely girls friend and the goal of trying to make her smile despite the stress she was under.
So soon did her zest for the rules crumble when she finally began to care about Marinette in a way she never had for anyone but her destructive other half. How could she insist that Marinette only use her powers to fight evil when swinging through the air on a dark Paris night gave her such joy? How could she insist that her duties as a Holder came first, when the poor girl hadn't seen her few friends in ages?
How could Tikki insist that Marinette must remain virtuous and righteous when that filthy false fox tried to ruin the few happy things her precious human had left?
I'm just....mystified and fascinated by the idea of a Tikki and Plagg that are more directly godlike and Eldritch as they've been mainly cut off from close contact and relationships with humans before, but who are also unable to stop themselves from becoming utterly enamored of their first, and possibly only, holders.
The other Kwami have had millennia to learn to let go, to let human things happen, and to stay out of their holders everyday conflicts. But Tikki and Plagg were likely alone in the miracle box so often before Fu's mistakes. And even though they've had their fellow Kwami for a human lifetime or two, they've still never truly experienced a partnership or friendship this close. I think the idea that they'd become over protective and borderline irrational about their chosen safety is a very likely outcome from all of this.
Also, I'm just a sucker for eldritch beings of immense power legitimately and honestly caring about tiny squishy humans, and very much near constantly overreacting about it.
Also, if fu tried to take their miraculous back in this worlds Feast, they might honestly just try to kill him. Damage to Paris from their actions be damned. They are not leaving their kids.
I also wanna explore like, Plagg urging Adrien to explore his more destructive desires(mostly healthily), and Tikki cultivating Marinette's creative and artistic talent. Just, them pushing their chosen holders to more deeply connect with their domains.